The medial entorhinal cortex keeps Up.

Dupret D
Csicsvari J
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He S, Mostofi A, Syed EC, Torrecillos F, Tinkhauser G, Fischer P, Pogosyan A, Hasegawa H, Li Y, Ashkan K, Pereira EA, Brown P, Tan H
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He S, Everest-Phillips C, Clouter A, Brown P, Tan H
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Guerra A, Pogosyan A, Nowak M, Tan H, Ferreri F, Di Lazzaro V, Brown P

2016.Cereb. Cortex, 26(10):3977-90.